Tver

Tver was a Rus' principality on the upper Volga, ruled from Tver by Yaroslav Yaroslavich's descendants. In 1337 Constantine ruled after the anti-Tatar uprising of 1327 had left Tver burned and Moscow in possession of the Vladimir title.

In 1375 a Muscovite siege forced Prince Mikhail II to acknowledge Dmitry Donskoy as his elder brother. Tver allied with Lithuania's Vytautas in 1427, but when Prince Mikhail III sought Lithuanian help, Ivan III conquered and annexed the principality in 1485, granting it to his son Ivan the Young.